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Shortly after Cole & Hart [1930] had reported the discovery of a gonadotrophic hormone in the blood of the mare in early pregnancy they [Cole, Howell & Hart, 1931] described the changes that occur in the ovaries throughout gestation, and associated the unusually active condition of the ovaries between the 40th and 150th day with the presence of the gonadotrophin in the blood. During this period the ovaries are very large and contain numerous follicles and a small number of functional corpora lutea. The authors claim that the latter serve initially to supplement the function of the primary corpus luteum, which even at this early stage of pregnancy is in decline, and ultimately to maintain the endometrial conditions requisite for the continuation of pregnancy.

Since the above observations were made, much has been learned regarding the site of origin [Cole & Goss, 1943; Rowlands, 1947] and the biological standardization of

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